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. 2017 Oct 26;7:14161. doi: 10.1038/s41598-017-14036-1

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Proxy measures estimating extent of path integration (A) and centre of search (B). (A) The maximum value along the y-axis, which runs from the release point (0) in the feeder-to-nest direction, in the first 10 m of path length. (B) The mean value of all digitised points along the y-axis in the segments of paths from 10 m to 20 m of path length. The top dotted line indicates the y value of the fictive nest, where the nest would lie had the ants not been displaced for the test. The bottom dotted line indicates the y value of the release point. The boxes show the median in the centre line, and interquartile ranges at their top and bottom. The tails show values that are 1.5 times the distance from the median to each quartile. The + symbols show outlying values beyond the tails. PI = path integration.