(A-H) BrdU was injected for 90 min to label proliferating cells in the liCL and laCL (denoted by white, dashed lines). Magnified view of the liCL (A′,B′) showed an increase in the number of proliferating cells in mutants (B′) compared to controls (A′). The number of BrdU+ cells was quantified and confirmed to be increased in mutants (C) however, the length of the BrdU+ zone was unchanged. Conversely, in the laCL, the number of BrdU+ cells was quantified and shown to be decreased in mutants compared to controls (E,F,G). Furthermore, the length of the BrdU+ zone was significantly shortened in mutants (H). **, P<0.01.