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The F. columnare type IX protein secretion system. Orthologs of proteins required for secretion in Flavobacterium johnsoniae and Porphyromonas gingivalis are shown in orange. F. johnsoniae PorV is required for secretion of many but not all proteins that are targeted to the T9SS. F. johnsoniae PorU is involved in but not essential for secretion. Black lines on the lipoproteins GldK and SprE indicate lipid tails. OM, outer membrane; CM, cytoplasmic membrane.
