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. 2017 Aug 24;8(55):93593–93607. doi: 10.18632/oncotarget.20547

Table 4. Associations of selected loci with leptin levels.

Gene Gene expression abundance in hypothalamus* SNP Effect size Leptin (ng/ml) [geometric mean (95% CI)] Padjusted1 Padjusted1+BMI
aa aA AA
PCSK1 32.4 (10.3) rs261967 0.4% 4.6 (4.2-5.0) 5.3 (5.0-5.7) 5.2 (4.7-5.8) 0.003 0.419
MC4R 3.8 (3.45) rs2331841 0.3% 4.7 (4.5-5.0) 5.5 (5.1-5.8) 5.6 (4.7-6.6) 0.004 0.694
FTO 543.45 (119.7) rs1558902 0.3% 4.9 (4.6-5.1) 5.7 (5.2-6.3) 6.0 (4.1-8.7) 0.008 0.216
MAP2K5 6.0 (5.6) rs4776970 0.3% 4.8 (4.5-5.1) 5.4 (5.0-5.8) 6.0(5.2-7.0) 0.013 0.151
GNPDA2 6.6 (6.65) rs16858082 0.2% 4.7 (4.4-5.1) 5.2 (4.9-5.6) 5.4 (4.8-6.2) 0.019 0.838
BDNF 12.4 (5.9) rs2030323 0.2% 4.8 (4.3-5.3) 4.9 (4.6-5.2) 5.5 (5.0-5.9) 0.053 0.235
GPSleptin / six leptin-related loci 1.6% Tertile1 (0-3 risk alleles) (n = 1373) Tertile2 (4 risk alleles) (n = 828) Tertile3 (5-9 risk alleles) (n = 1130) 2.8E-6 0.218
4.4 (4.1-4.8) 5.1 (4.7-5.6) 5.8 (5.4-6.3)

Abbreviation: PCSK1, proprotein convertase subtilisin/kexin type 1; MC4R, melanocortin 4 receptor; FTO, fat mass and obesity associated; MAP2K5, mitogen-activated protein kinase 5; GNPDA2, glucosamine-6-phosphate deaminase 2; BDNF, brain-derived neurotrophic factor and GPSleptin, sum gene score of six leptin-related loci.

*Data were expressed as mean (median in all tissues and cells), from http://biogps.org.

Padjusted1 means the analysis of covariance in the general linear model after adjusting for gender, age, Tanner stage and residence.

Padjusted1+BMI: further adjusted for BMI.

Note: Values in bold are significant at P < 0.05.