Changes in orientation preference and receptive field position along a horizontal track of cat visual cortex. From top to bottom, the rows show OFF receptive fields mapped with dark stimuli (in blue), ON receptive fields mapped with light stimuli (in red), the ON-OFF receptive field difference (diff.), and the orientation/direction preference measured with moving bars (circles show the orientation preference predicted from the receptive field maps). From left to right, the columns show cortical measurements separated by 0.1 mm from 0 to 1.2 mm distance. The column on the right shows the receptive field average along the entire horizontal track (top three receptive fields) and the central positions of ON (red circles) and OFF receptive fields (blue circles). The position of these ON and OFF cortical receptive fields is determined by the receptive field population of the ON and OFF thalamic afferents.