Caves considered in this study. Numbers between brackets correspond with the caves shown in Figure 1 and Figure 3. Water presence has four ordered levels (1, occasional presence of areas of water or humid spots; 2, presence of scattered areas of water in the form of more or less continuous drops; 3, presence of permanent pools throughout the cave; 4, presence of rivers or streams), the tropic level has three ordered levels (1, oligotrophic, not having organic material; 2, mesotrophic, scattered organic material present along the cave; 3, meso‐eutrophic, with accumulations of organic material present along the cave), and genesis has two categories (0, epigenic; 1, hypogenic)