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. 2017 Dec 19;7:17791. doi: 10.1038/s41598-017-18104-4

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© The Author(s) 2017

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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(A) Grooming networks constructed using matrices of directional rates of grooming. Social group and year of observation are indicated under each network. Males are squares, females circles. High ranking animals are darker colours. (B) Heatmaps of network measures across years for individuals with >2 years of data: (B1) instrength, (B2) outstrength, (B3) clustering coefficient, (B4) betweenness, (B5) eigenvector centrality. Network measures range from low values (yellow) to high values (red) as shown in legend bars. Blank cells represent years with no data collected for an individual, grey cells in B3 represent animals with one or few social partners and thus null clustering coefficients.