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His516 side chain dihedral angles (χ1 and χ2) distribution: (a) WT GOx, (b) parent mutant P, (c) Pk, (d) Pv, (e) A2, and (f) F9. The χ1 dihedral has a clear preference for g– geometry. The χ2 dihedral prefers either the catalytic Nt or noncatalytic Ng+ geometries in the different GOx variants. The normalized integrated distributions are shown as red curves on either side of the panels.
