Fig. 4.

Diversification patterns of major beetle lineages inferred from MEDUSA analysis. Each terminal represents a monophyletic family-level taxon. The number in parenthesis next to the taxon name indicates the number of validly described species within the taxon. The species richness of each taxon is also indicated with histograms on the right. Branches are color coded to show the diversification rates. Clades with significant diversification rate shifts compared with the background rate are marked with circled numbers on the tree (red: rate increase; blue: rate decrease). Estimated net diversification rates and differences in AICc scores are included in the lower left table