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. 2018 Jan 19;6:e4279. doi: 10.7717/peerj.4279

Table 2. Midshaft humeral unstandardized properties of Zhoukoudian left humerus (II) and comparative samples.

Length Body mass TA CA %CA Imax Imin Zmax Zmin J Zp
(mm) (kg) (mm2) (mm2) (mm4) (mm4) (mm3) (mm3) (mm4) (mm3)
Zhoukoudian IIa,b 307.4–324.0 53.6 ± 1.7 261 228 87.4 6,985 4,143 640 518 11,128 1,009
Tianyuan 1a,c 327.4 85.1 252 190 75.4 5,931 3,868 603 463 9,799 928
Middle Paleolithic Modern Humand (n = 2 for length, body mass, TA, CA, Ix, and Iy, n = 3 for Imax, Imin, and J) Mean 353.3 68.9 283.1 217.0 76.8 5,894 4,088 9,981
S.D. 30.8 0.1 5.2 56.9 21.5 2,021 1,619 3,618
Min 331.5 68.8 279.4 176.7 61.6 3564 2,287 5,851
Max 375.0 69.0 286.7 257.2 92.1 7,170 5,421 12,591
Neanderthald (n = 5 for length, n = 4 for body mass, n = 7 for TA and CA, n = 6 for Ix and Iy, n = 8 for Imax and Imin, n = 9 for J) Mean 314.4 79.1 256.0 197.8 77.6 7,879 4,173 12,112
S.D. 13.4 9.7 44.0 29.3 3.6 2,863 1,658 4,199
Min 299 64.8 203.5 170.7 73.9 4,629 2,250 6,879
Max 334 85.5 341.1 251.9 84.2 12,020 6,411 18,250
Early Upper Paleolithic Modern Humand (n = 20 for length, n = 15 for body mass, n = 17 for TA, CA, Ix, and Iy, n = 22 for Imax and Imin, n = 23 for J) Mean 326.5 68.4 298.6 198.6 67.1 7,119 4,799 12,138
S.D. 21.0 7.7 46.1 29.5 8.9 1,965 1,315 2,978
Min 288.0 54.3 199.8 133.0 47.9 3,670 2,148 5,895
Max 370.0 82.5 394.1 246.7 83.0 10,701 7,316 17,605
East Eurasia Late Upper Paleolithic Modern Humand (n = 7 for length, n = 5 for body mass, n = 10 for TA, CA, Ix, Iy, Imax, Imin, and J) Mean 273.1 53.2 227.6 168.4 74.2 5,106 2,972 8,078
S.D. 20.3 10.5 33.8 27.9 7.6 1463 955 2,395
Min 250.0 42.3 186.7 138.8 65.7 3,437 1,900 5,587
Max 311.0 70.5 281.8 225.1 86.5 7,432 4,724 11,968

Notes.

a

Estimated cross section location due to incomplete length.

b

Maximum length of the left Zhoukoudian Humerus II was reported by Weidenreich (1941) to be 324.0 mm. We estimated maximum length as 307.4 mm using a regression analysis of the distance between the deltoid tuberosity and the proximal margin of the olecranon fossa against maximum length on our comparative sample of Datong and Junziqing modern Homo sapiens (n = 33; see Text S4). In order to be conservative, we use both estimates to provide a range of standardized values for Zhoukoudian humeri about a mean value (315.7 mm). We estimated cross-sectional properties of Humerus II from its periosteal contour, and a radiograph published by Weidenreich (1941: Fig. 58B); see Text S3.