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. 2018 Jan 22;8:1300. doi: 10.1038/s41598-017-14403-y

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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(A) The read-pair types at the boundary between telomere and subtelomere. F2a reads stem from the boundary whereas F1 reads stem from anywhere within the telomere proper. F3 are reads where neither read in the pair is complete telomere (B) Detail of the F1 and F2a read types. F1 read-pairs are comprised of two complete telomere reads. F2a read-pairs are comprised of a read-pair where one read is complete telomere and the other is not. Crucially, the complete telomere read is comprised of CCCTAA (C) The read-pair types at an ITR. (D) Detail of the F2b and F4 read types. Note that the F2b is physical indistinguishable from an F2a read. An F4 read is read-pair where one read is complete telomere and the other is not. The complete end is comprised of TTAGGG.