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. 2018 Jan 23;8:1409. doi: 10.1038/s41598-018-19697-0

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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(A) Specific biofilm unit of wild-type H. pylori J99, J99ΔluxS and J99ΔcagY determined by crystal violet assay. (B) Comparison of number of metabolites detected by LC-MS per sample for H. pylori J99, J99ΔluxS and J99ΔcagY. *Represents p-value < 0.05 comparing between wild-type and mutants. (C) Comparing the relative abundance of PI-Cer(t18:0/18:0(2OH)) in H. pylori J99, J99ΔluxS, J99ΔcagY, high biofilm-forming clinical strains and low biofilm-forming strains.