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. Author manuscript; available in PMC: 2019 Mar 1.
Published in final edited form as: J Math Biol. 2017 Jul 24;76(4):911–944. doi: 10.1007/s00285-017-1160-3
j Competition process Birth/birth-death

(a + 1, b) q(a,b)(a+1,b)
λab(1)
(a − 1, b) q(a,b)(a−1,b) 0
(a, b + 1) q(a,b)(a,b+1)
λab(2)
(a, b − 1) q(a,b)(a,b−1)
μab(2)
(a + 1, b − 1) q(a,b)(a+1,b−1) γab
(a − 1, b + 1) q(a,b)(a−1,b+1) 0
(a, b)
-k,l{-1,0,1}klq(a,b)(a+k,b+l)
-(λab(1)+λab(2)+μab(2)+γab)
other 0 0