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. 2018 Feb 1;6:e4277. doi: 10.7717/peerj.4277
1. Antenna 12-segmented 2
–Antenna 11-segmented 3
2. Body pilosity clubbed; mesosoma, petiole and postpetiole distinctly sculptured (Fig. 4A) sarawatense Sharaf & Aldawood
–Body pilosity simple; mesosoma, petiole and postpetiole smooth and shining (Fig. 4B) holothir Bolton
3. Mesosoma without standing hairs (Figs. 4C and 4D) 4
–Mesosoma with standing hairs (Figs. 4E and 4F) 5
4. Eyes appearing larger, with a ring of seven to eight ommatidia encircling a single row of 2 ommatidia, and in profile closer to mandibular insertions (EM 0.05); meso-and metapleuron smooth; petiole and postpetiole smooth and each with one pair of standing hairs (Fig. 4C) aeyade Collingwood & Agosti
–Eyes appearing smaller, with only 5–6 ommatidia, and in profile further away from mandibular insertions (EM 0.09–0.11); meso-and metapleuron finely shagreened; petiole and postpetiole superficially shagreened and without standing hairs (Fig. 4D) mohammedi sp. n.
5. Mesosoma with only two pairs of standing hairs, one on pronotal corners and one propodeum (Fig. 4E) clavicorne André
–Mesosoma with several pairs of standing hairs, about 10 pairs (Fig. 4F) exiguum Forel