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The eukaryotic replisome positions Pols ε and α on opposite sides of CMG. Depending on whether the CTD motor ring or the NTD ring pushes against the forked junction, the polymerases are positioned either: (a) with Pol ε in front and Pol α-primase in back of CMG or (b) with Pol α-primase in front and Pol ε in back of CMG
