Fig. 4.

Fitness effects of the deletion and A301S alleles measured in 20,000-generation clones isolated from each population. If the evolved allele present in a clone was different from either of the transferred alleles, then its effect is also shown as the “original” allele. If an evolved clone already had one of the transferred alleles, then this fact is indicated by red fill in the corresponding symbol for the deletion (for Ara–1) or A301S (for Ara–5, Ara+1, and Ara+5) allele. Error bars indicate 95% confidence intervals (n = 3).