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Multiple sequence alignment of the CE4 enzymes listed in Table 1. Loops are highlighted with colored boxes according to [62]. Conserved catalytic motifs are labelled MT1–5. The “His–His–Asp” metal binding triad (▼), catalytic base (*), and catalytic acid (◊) are labelled. The mark inside Loop 5 for poly-β-1,6-GlcNAc deacetylases (four last sequences) indicates the shuffling point of the circularly permuted CE4 domain.
