Levels of organic acids in the seedlings of AtUCP knockouts exposed or not to salt stress. Plants were grown on the plates containing 0, 50, and 75 mm NaCl for 12 days, and the relative levels of malate (A), fumarate (B), citrate (C), and pyruvate (D) in whole seedlings were determined. The levels of the metabolites were normalized to the mean of those of wildtype plants grown on the plate without salt. The means ± S.E. (error bars) from plants grown on three individual plates are shown. Orange, wildtype; brown, ucp1; green, ucp2; light green, ucp1/ucp2 double knockout.