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a) CMG alternates between the compact and extended conformers (PDB 3JC5, and 3JC7). b) Inchworm process proposed for movements between the N- tier and C-tier (N-tier in solid red, because it directly contributes to translocation by binding DNA). c) Inchworm process proposed for planar to spiral movments solely in the C-tier (N-tier shown in light blue, because it is a processivitiy ring and does not participate in inchworm translocation).
