Table 1.

Principal components of flavonoids and phenolic compounds of different oregano species.

Oregano Species	Origin	Extraction Solvent	Flavonoids and Phenolic Acids Constituents	Reference
Hedeoma patens	Mexico	Methanol/acetone/water (50:40:10)	3-O-caffeoylquinic acid, luteolin-7-O-glucuronide, scutellarein-7-O-hexoside, salvianolic acid A	[4]
Lippia alba (Mill.) N. E. Brown	Brazil	Methanol 62.5%	Apigenin-7-O-diglucuronide, chrysoeriol-7-O-diglucuronide, luteolin-7-O-glucuronide, tricin-7-O-diglucuronide, tricin-7-O-glucuronide	[25]
Lippia alba (Mill.) N. E. Brown	France	Methanol/hydromethanol	Luteolin-7-diglucuronide, verbascoside, chlorogenic acid	[28]
Lippia alba (Mill.) N.E. Brown	Brazil	Chloroform	5,5′′-dihydroxy-6,4′,6′′,3′′′,4′′′-pentamethoxy-[C7–O–C7′′]-biflavone, 4′,4,5,5′′-tetrahydroxy-6,6′′,3′′′-trimethoxy-[C7–O–C7′′]-biflavone	[22]
Lippia citriodora	Germany	Water	Verbasoside, luteolin-7-diglucuronide, apigenin-7-diglucuronide	[29]
Lippia graveolens	Mexico	Methanol/acetone/water (50:40:10)	Quercetin O-hexoside, scutellarein 7-O-hexoside, phloridzin, trihydroxy-methoxyflavone derivative, 6-O-methylscutellarein	[4]
Lippia graveolens	NR 1	Methanol 70%	6-Hydroxyluteolin 7-O-hexoside, pentaflavanone-B hexoside-1, pentaflavanone-B hexoside-2, scutellarein 7-O-hexoside, luteolin 7-O-glucoside, taxifolin, 6-hydroxyluteolin 7-O-rhamnoside, 6-hydroxyluteolin, 3-hydroxyphloretin 6′-O-hexoside, apigenin 7-O-glucoside, phloridzin, scutellarein, eriodictyol, luteolin, quercetin, naringenin, 6-methylscutellarein, 6,7-dimethylscutellarein, sakuranetin, pinocembrin, galangin, methylgalangin	[9]
Lippia graveolens	USA	Methanol 100%	Eriodictyol, naringenin, hispidulin, cirsimaritin	[30]
Lippia micromera	Colombia	Methanol/hydromethanol	Naringenin, apigenin	[28]
Lippia origanoides	Colombia	Methanol 62.5%	Quercetin, naringenina, luteolin, pinocembrin	[25]
Lippia palmeri	Mexico	Methanol/acetone/water (50:40:10)	Quercetin-O-hexoside, luteolin 7-O-glucuronide-3′-O-glucoside, scutellarein 7-O-hexoside, trihydroxy-methoxyflavone derivative	[4]
Majorana hortensis Moench.	Poland	Water	Caffeic acid, lithospermic acid, rosmarinic acid	[31]
O. acutidens	Turkey	Water	Gallic acid, caffeic acid, 4-hydroxybenzaldehyde, p-coumaric acid, rosmarinic acid	[32]
O. dictamus	Greece	Methanol 62.5%/BHT	Vanillic acid, protocatechuic acid, syringic acid, gallic acid, cinnamic acid, o-coumaric acid, p-coumaric acid, caffeic acid, chlorogenic acid, rosmarinic acid, chrysin, epicatechin, naringenin, catechin, genistein, quercetin	[33]
O. dictamus	Greece	Water	Chlorogenic acid, rutin, luteolin-7-O-glucoside, apigenin-7-O-glucoisde, rosmarinic acid, luteolin	[34]
O. indercedens	Greece	Methanol	Caffeic acid, rosmarinic acid	[35]
O. majoram L.	Germany	Methanol 60%/Formic acid 1%	Apigenin-6,8-di-C-glucoside, luteolin-7′-O-glucuronide, rosmarinic acid, apigenin-glucuronide, lithospermic acid A isomer a, salvianolic acid B, apigenin	[36]
O. majoram L.	Tunisia	Methanol	Dihydroxybenzoic acid hexose, syringic acid, vanillic acid, dihydroxybenzoic acid, p-coumaric acid, chlorogenic acid, salvianolic acid I, caffeoyl-arbutin, rosmarinic acid, gallocatechin isomer 1, gallocatechin isomer 2, 3-O-methyl-catechin, luteolin-6,8-C-dihexose, apigenin-6,8-di-C-hexoside, isoorientin, orientin, isovitexin, kaempferol-O-sumbubioside, kaempferol-O-glucuronide, luteolin-O-glycoside, diosmin, apigenin-O-glucuronide, acacetin rutinoside, luteolin, apigenin, taxifolin, taxifolin methyl ether isomer 1, taxifolin methyl ether isomer 2, dihydrokaempferide, hesperidin, eriodictyol, sakuranetin, O-methy-quercetin, dimethyl myriceitn, quercetin dimethyl ether, jaceidin isomer 1, jaceidin isomer 2	[37]
	Greece	Water	Vanillic acid, protocatechuic acid, syringic acid, gallic acid, cinnamic acid, o-coumaric acid, p-coumaric acid, ferulic acid, caffeic acid, sinapic acid, rosmarinic acid, chrysin, epicatechin, naringenin, catechin, kaempferol, quercetin	[34]
	Poland	Methanol	Protocatechuic acid, p-hydroxybenzoic acid, gentisic acid, chlorogenic acid, syringic acid	[38]
	Turkey	Methanol 80%	Caffeic acid glucoside, epigallocatechin, arbutin, luteolin ruitinoside, luteolin glucuronide, rosmarinic acid, dihydroquercetin, dihydroluteolin, apigenin, quercetin, quercetin arabinoside, luteolin-7-O-glucoside gallocatechin derivative	[39]
	Germany	Methanol 50%	Luteolin-6,8-di-C-glucoside, apigenin-6,8-di-C-glucoside, luteolin-glucuronide, rosmarinic acid, apigenin-glucuronide, lithospermic acid, apigenin	[40]
O. majoram L.	USA	Methanol 80%	Eriodictyol 6,8-di-C-glucoside, eriodictyol 7-O-glucoside, apigenin 6,8-di-C-glucoside, luteolin 7,7′-di-O-glucuronide, luteolin 7-O-glucuronide-3′-O-glucoside, apigenin 7-O-diglucuronide, luteolin 7-O-glucuronide, luteolin 7-O-glucuronide, luteolin 7-O-glucoside, apigenin 7-O-glucoside, apigenin 7-O-glucuronide, rosmarinic acid	[41]
	Turkey	Water	Gallic acid, caffeic acid, p-coumaric acid, rosmarinic acid, chicoric acid, apigenin-7-glucoside, quercetin, kaempferol	[32]
	Italy	Cascade extraction with ethyl acetate and ethanol	Eriodictyol 7-O-rutinoside, aromadendrin, eriodictyol, naringenin, luteolin, sorbifolin, cirsiliol, apigenin, cirsilineol, cirsimaritin, xanthomicrol, caffeic acid, rosmarinic acid	[42]
	Greece	Cascade extraction with hexane and ethyl acetate	Salvianolic acid H, salvianolic acid B, rosmarinic acid, salvianolic acid C, eriodictyol, naringenin	[43]
	Finland	Methanol 40%	Calleryanin 3,4-dihydroxybenzoate, gastrodin 3,4-dihydroxybenzoate, calleryanin 3-hydroxy,4-methoxybenzoate	[44]
	Portugal	Methanol 80%	Gallic acid, 3,4-dihydroxybenzoic acid, (+)-catechin, caffeic acid, (−)-epicatechin, rosmarinic acid	[10]
Oregano 2	Turkey	Methanol 80%	Gallic acid, syringic acid, vanillic acid, protocatechuic acid, chlorogenic acid, p-coumaric acid, quercetin-3-O-hexoside, luteolin-7-O-glucoside, ferulic acid, phloridzin, dicaffeoylquinic acid, apigenin-7-O-rutinoside, rutin, apigenin-7-O-glucoside, rosmarinic acid, luteolin-3-O-glucuronide, luteolin-7-O-rutinoside, quercetin, apigenin, 4′-methoxyapigenin, 6,7-dimethoxyscutellarein, luteolin	[45]
Poliomintha longiflora	NR 1	Phosphate buffer	Vanillic acid, caffeic acid, luteolin, rosmarinic acid, hispidulin	[46]

¹ Not reported, ² Species not mentioned.