Table 3.

The effect of shading and drought on dark respiration (R_dark), maximum CO₂ assimilation (A_max), photosynthetic rate under saturating light (A_sat), and efficiency of PSII (Φ_PSII) of the four study species. Means (±SE) include 16 replicates per treatment, and means not connected by the same letter are significantly different (Tukey HSD, p<0.05).

Treatment	Rdark (μmol CO2 m−2 s−1)	Amax (μmol CO2 m−2 s−1)	Asat (μmol CO2 m−2 s−1)	ΦPSII (μmol CO2 μmol−1 photons)
Control	1.3±0.1 a	35.4±3.2 a,b	19.3±2.7 a	0.152±0.015 a
Drought	1.1±0.1 a	37.4±3.5 a	13.1±1.7 b	0.140±0.012 a
Shade	0.9±0.1 b	31.4±2.9 b	16.5±2.2 a,b	0.108±0.011 b
Shade + Drought	0.8±0.1 b	32.6±2.4 a,b	15.3±2.0 a,b	0.111±0.010 b