Expansin-induced primordia develop into normal leaves and lead to
reversed phyllotaxy. (A) Expansin-induced leaf.
(B) Normally formed leaf. (C) Cross
section of the midrib of an expansin-induced leaf. (D)
Cross section of the lamina of an expansin-induced leaf.
(E and F) Reversal of phyllotaxy after
the initiation of a leaf (I′2) (arrow in E) by localized
induction of expansin expression. As shown in diagram
(F), initially leaves (P3, P2, P1) were formed in an
anticlockwise direction. Leaves formed subsequent to I′2 (I′3- I′7)
have been formed in a clockwise direction. (G) Side view
of plant shown in E to demonstrate leaf insertion points
along the stem. e, epidermis; p, palisade; s, spongy mesophyl. (Bar:
A and B= 5 mm; C and
D = 125 μm.)