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. 2018 Apr 11;8:5802. doi: 10.1038/s41598-018-24223-3

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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A structural model of mucus comprising rheologically heterogeneous domains blended together. (A) Periodic acid-Schiff staining of the pseudostratified columnar epithelium from the proximal small intestinal porcine mucosa which illustrates the presence of domains in physiological mucus. Note the pink staining of mucus from goblet cells into the intestinal lumen and the blue staining of nuclei. Scale bar 20 μm. (B) Visualization of the rheological heterogeneity, X- and Y-positions, of movement of individual 0.5 μm colloidal particles in 50 mg/mL ND-mucin solution. The size of circles indicates diffusivity and the color corresponds to the power law scaling of the MSD spectra.