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. 2018 Apr 11;9:1385. doi: 10.1038/s41467-018-03783-y

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Fig. 5 — The effect of physical properties on the apparent fitness landscape (AFL). Each value represents the sum of the apparent fitness score (AFS) multiplied by a scalar corresponding to the indicated property. Blue scores mean that the position showed a preference for high property values, while red scores show a preference against high property values. Mutability index is shown as a reference