Distribution of peltate gland developmental stages among leaves of various ages. Bars indicate the percentage of the total gland number represented by each developmental stage (left axis). The line graph indicates the total number of peltate glands (right axis) for each leaf size class. Peltate gland stages: 2, two-celled apical disc; 4, four-celled; 8s, eight-celled, small; 8L, eight-celled, large; S−, early secretory stage; S, middle secretory stage; S+, post-secretory stage. Error bars represent ses.