Model depicting the function of RMD in rice pollen tube growth. RMD, present at the tip of the pollen tube, functions as a key regulator that contributes to the F-actin-dependent maintenance of the spatial organization of cytoplasmic streaming, which is essential for pectin distribution to determine the tip growth of the rice pollen tube. Gray arrows trace the reverse-fountain cytoplasmic streaming pattern. DM, demethyl-esterified; M, methyl-esterified.