Heterologous expression of AtBBX21 increases anthocyanin and phenolic levels in potato leaves. A, Phenolics and anthocyanins in nontransformed (Spunta) and transgenic (CH1, CH2, and CH13) potato leaves (n = 5). B, Sunscreen pigment accumulation in young and totally expanded leaves. Sunscreen accumulation is revealed by a decrease in the intensity of UV-induced chlorophyll fluorescence (less fluorescence indicates higher accumulation of anthocyanin and phenolic compounds). C, Maximal photochemical efficiency (Fv/Fm) in 14-d-old plants acclimated at low irradiance (200 μmol m−2 s−1) and then exposed at high irradiance (900 μmol m−2 s−1) for 2 h (n = 6). n = number of biological replicates. Mean values are shown with error bars indicating se. Data were analyzed by Student’s t tests, and asterisks indicate significant difference between nontransformed and transgenic lines (*P < 0.05 and **P < 0.01; NS, not significant).