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. 2018 May 4;8:7039. doi: 10.1038/s41598-018-25468-8

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Deletion of IP6K1 elicited disorientation and malformation of elongating/elongated spermatids. (a–f) H&E staining of WT and IP6K1 KO testes. Stages of seminiferous tubules were determined by spermatid developmental status. (a,b,c,d,f) Cell polarity and morphology of the step 12–16 elongating/elongated spermatids were disrupted in IP6K1 KOs (insets). Scale bar 20 μm.