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. 2018 May 2;(98):51–71. doi: 10.3897/phytokeys.98.25044
1 Leaf blade transverse-cordiform in outline, pedately compound 2
Leaf blade cordiform-sagittate, sagittate or hastate in outline, margins entire, repand, sinuately lobed, pinnatifid or bipinnatifid 3
2 Ovary locules 3–4; leaflets 8–11, central leaflet 10–17 cm long; occurring on rocks in semi-arid areas or terrestrial in coastal restinga scrub on sand; usually in association with populations of Bromeliaceae; northeast Brazil T. leal-costae
Ovary locules 10–26; leaflets 10–20, central leaflet 18–50 cm long; hemiepiphytic or terrestrial; most common along river margins; Amazon basin T. spruceanum
3 Leaf margin sinuately lobed, pinnatifid or bipinnatifid 4
Leaf margin entire or repand or, if sinuately lobed, then peduncle 16 cm long or more 11
4 Leaf margin bipinnatifid, rarely pinnatifid but then with primary lateral veins of anterior division (5-)6–9(-10) per side; leaf blade over 50 cm long, primary lateral lobes (12-)17–35(-55) cm long 5
Leaf margin undulate or pinnatifid; if pinnatifid, then with primary lateral veins of anterior division 3–4(-10) per side; leaf blade up to 50 cm long, usually smaller; primary lateral lobes 5–17.5 cm long 7
5 Petioles flattened or slightly convex adaxially; intravaginal squamules never persistent, foliage leaf scars always concolorous with the internodes; uplands of Cerrado (Minas Gerais, Bahia and Goiás states – 700–1200 m) T. lundii
Petioles conspicuously sulcate adaxially; intravaginal squamules persistent, very rarely deciduous (if deciduous, then foliage leaf scars discolorous with the internodes) 6
6 Intravaginal squamules numerous and dense, 5–12 mm long, 2–4 mm wide at base, persistent but easily detachable, rarely deciduous; female portion of the spadix adnate to the spathe for 60–80% of its length; southern and western (coastal) Brazil, Argentina, Paraguay T. bipinnatifidum
Intravaginal squamules few and scattered, robust, 8–20 x 5–10 mm, always persistent, hardly detachable; female portion of the spadix adnate to spathe for 40–50% of its length; northern Goiás and possibly Mato Grosso states T. mello-barretoanum
7 Plants aquatic or rarely terrestrial; leaf margin sinuately lobed (sinuses penetrating less than halfway to midrib), primary lateral lobes of anterior division 1.5–6.5(-14) cm long, usually oblique and turned towards leaf apex; female zone of the spadix (1.5-)4–5 cm long 8
Plant rupiculous or terrestrial; leaf margin pinnatifid (sinuses penetrating at least halfway to midrib), primary lateral lobes of anterior division 5–17.5 cm long, not oblique, female zone of the spadix 1.4–3.4 cm long 9
8 Stems with long and thorn-like intravaginal squamules; leaf blades never erect in living plants; Argentina, Paraguay, Bolivia and Brazil (South and South-eastern) T. undulatum
Stems without persistent intravaginal squamules; leaves always erect or semi-erect in living plants; Eastern Brazil (northern Espírito Santo and southern Bahia) T. paludicola
9 Petiole green or glaucous green at apex; leaf blade broadly ovate in outline, dark to subglossy green, sometimes glaucous, primary lateral lobes 3–4(-5); spathe 6.4–16 cm long, green outside, opened at anthesis; ovary locules 4–8(-11) 10
Petiole purplish at apex; leaf blade triangular to ovate in outline, glossy dark green, primary lateral lobes 5–10; spathe (8.2-)12–18 cm long, dark purple outside, tightly clasped around spadix at anthesis; ovary locules (6-)7–8 T. xanadu
10 Leaf blade (32-)35–50 cm long, primary lateral lobes of anterior division 3.5–6.3(-7.5) cm wide, distance between sinuses and midrib progressively greater towards base of anterior division; fertile male zone of the spadix 1.5–2.2 cm diam. T. saxicola
Leaf blade 17-33 cm long, primary lateral lobes of anterior division 1.4–3.7(-7.4) cm wide, distance between sinuses and midrib usually becoming progressively less towards base of anterior division; fertile male zone of spadix 0.85–1.3 cm diam. T. adamantinum
11 Overall length of adult leaf blade more than 60 cm (sometimes 50–60 cm in T. solimoesense); petiole apex often minutely rugose-verruculate (may be smooth in T. stenolobum) 12
Overall length of leaf blade less than 60 cm, petiole apex smooth, never occurring in Amazonia 16
12 Species from Eastern Brazil; stamens 6 mm long or more; staminodes more than 1.6 mm wide at apex, less than 2.5× longer than wide; ovary locules 6–13 per ovary 13
Species from Amazonia; stamens less than 6 mm long; staminodes less than 1.6 mm wide at apex, more than 2.5× longer than wide; ovary locules 17–34(-47) per ovary 15
13 Leaf blade narrowly sagittate, sometimes subhastate; anterior division 2.1–3.3× longer than wide; intravaginal squamules deciduous T. stenolobum
Leaf blade broadly sagittate; anterior division 1–1.5× longer than wide; intravaginal squamules small but persistent 14
14 Leaf blade less than twice as long as wide; spathe outside lacking extrafloral nectaries, inside carmine magenta at anthesis; central style dome lacking T. speciosum
Leaf blade more than twice as long as wide; spathe outside with punctate, pale brown extrafloral nectaries, inside cream-white at anthesis; central style dome present T. williamsii
15 Cataphylls persistent; primary lateral veins of anterior division of leaf blade (5-)6–7; ovary locules 17–22; style elongated, distinctly narrower than ovary and lacking an axial canal T. venezuelense
Cataphylls deciduous; primary lateral veins of anterior division of leaf blade (3-)4–5(-6); ovary locules 26–34(-47); style short, as broad as ovary with an axial canal or cavity which is very conspicuous in fruit T. solimoesense
16 Peduncle subequal to twice as long as spathe; plants aquatic or rupiculous, aerial portion of the stem unbranched; internodes shorter than prophyll scars 17
Peduncle only about one third of spathe length; plant hemi-epiphytic or terrestrial; aerial stem branching frequently; internodes usually longer than prophyll scars T. corcovadense
17 Leaf blade at least twice as long as broad; style longer than ovary T. dardanianum
Leaf blade much less than twice as long as broad; style shorter than ovary 18
18 Intravaginal squamules abundant; broadly triangular, 3–12 mm long, (1.5-)3–7(-9) mm broad at base; style body as wide as ovary T. brasiliense
Intravaginal squamules few, more narrowly triangular, 1.5–5 mm long, 0.5–2.5 mm broad at base; style body slightly narrower than ovary 19
19 Leaf blades subglossy to glaucous green, margins weakly repand; Argentina, Paraguay, Uruguay, South Brazil 20
Leaf blades dark glossy green, margins entire, rarely repand; central Brazil T. uliginosum
20 Plants rupiculous, stem erect; prophyll deciduous; stylar central dome present T. petraeum
Plants aquatic, stem decumbent or rhizomatous subterranean; prophyll marcescent and persistent; stylar central dome absent T. tweedieanum