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. Author manuscript; available in PMC: 2018 Jun 1.
Published in final edited form as: Ecol Lett. 2018 Feb 28;21(6):763–778. doi: 10.1111/ele.12928
Source of covariance Sum of cross products
Total kPjTiN(ΔpijkΔp¯¯¯)(MijkM¯¯¯)=                    (E9)
Average selection effect PTiN(Δpi¯¯Δp¯¯¯)(Mi¯¯M¯¯¯)+
Temporal insurance effect PjTiN(Δpij¯Δpi¯¯)(Mij¯Mi¯¯)+
Spatial insurance effect TkPiN(Δpik¯Δpi¯¯)(Mik¯Mi¯¯)+
Spatiotemporal insurance effect kPjTiN(ΔpijkΔpij¯Δpik¯+Δpi¯¯+Δp¯¯¯)(MijkMij¯Mik¯+Mi¯¯+M¯¯¯)