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. 2018 May 15;(758):75–113. doi: 10.3897/zookeys.758.23019
1 Fore wings with distinct darker patches or clouds of pigmentation, in some cases only around termination of veins at wing margins and around cross veins between Rs and wing margin in cell r1 (Fig. 1E–I) 2
Fore wings without distinct darker patches or clouds of pigmentation, appearing uniformly clear or opaque, cross veins between Rs and wing margin in cell r1, if present, unpigmented (Fig. 1A–D) 6
2 Antennae short, subequal to head width (ratio AL:HW < 1.1), ratio HW:GP > 1.65, female proctiger strongly convex apically with apex extremely broad and blunt, ratio FP:HW < 0.70, on Hawaii 3
Antennae longer (AL:HW > 1.1), ratio HW:GP < 1.65, female proctiger more or less straight dorsally with apex bluntly acute, on other islands 4
3 Fore wings with extensive patches and clouds of red-brown pigmentation, particularly across the central area of wing, and numerous cross veins (typically more than 5) between Rs and wing margin in cell r1, smaller species with broader wings (ratio WL:WW < 3.1) and wing cell cu1 relatively narrow and high (ratio CUR < 1.33) (Figs 1I, 8A–W) S. rubra sp. n.
Fore wings with only indistinct fuscous brown on membrane and distinct small darker brown patches around termination of veins at wing margins and around cross veins between Rs and wing margin in cell r1, cross veins fewer (typically less than 5), larger species with narrower wings (ratio WL:WW > 3.1) and wing cell cu1 relatively wide and low (ratio CUR > 1.33) (Figs 1E, 5I–K) S. hawaiiensis sp. n.
4 Larger species (WL > 1.9 mm), paramere shape more triangular with a broader base, on Kauai (possibly other islands for S. reticulata) 5
Smaller species (WL < 1.9 mm), paramere shape more slender with a narrower base, on Oahu (Figs 1F, 11) S. magnaccai sp. n.
5 Fore wing with fewer cross veins (typically less than 6) between Rs and wing margin, head with shorter genal processes (ratio HW:GP > 1.36) and shorter antennae (AL < 0.75 mm, ratio AL:HW ≤ 1.40), paramere shorter and broader (ratio PL:HW < 0.20, ratio PL:AEL < 0.80), female terminalia shorter (ratio FP:HW < 0.80) with posterior margin of anal ring less convoluted and incised (Figs 1G, 9A–T; immatures Figs 12A–G, 13A–F) S. reticulata Caldwell, 1940
Fore wing with more cross veins (typically more than 6) between Rs and wing margin, head with longer genal processes (ratio HW:GP < 1.36) and longer antennae (AL > 0.75 mm, ratio AL:HW ≥ 1.40), paramere longer and narrower (ratio PL:HW > 0.20, ratio PL:AEL > 0.80), female terminalia longer (ratio FP:HW > 0.80) with posterior margin of anal ring more convoluted and incised (Figs 1H, 10A–S; immatures Figs 12H–N, 13G–J) S. tentaculata sp. n.
6 Smaller species (WL < 3 mm, AL < 1 mm), wing cell cu1 relatively narrow and high (ratio CUR < 2.1), on Oahu, Molokai, Maui 7
Larger species (WL > 3 mm, AL > 1 mm), wing cell cu1 relatively wide and low (ratio CUR > 2.1), on Kauai (Figs 1D, 6A–K) S. magna sp. n.
7 Genal processes shorter (GP < 0.40 mm, ratio HW:GP > 1.35), paramere shorter (ratio PL:HW < 0.27) and with less extended anteriorly directed apex, on Oahu 8
Genal processes longer (GP > 0.40 mm, ratio HW:GP < 1.35), paramere longer (ratio PL:HW > 0.27) and with more extended anteriorly directed apex, on Molokai and Maui (Figs 1A, 3A–P) S. elongagena Caldwell, 1940
8 Paler species (generally yellow-brown to green), fore wing vein Rs typically longer relative to wing length (ratio WL:Rs 1.57–1.82) resulting in shorter distance between terminations of Rs and M1+2, antennae longer (AL > 0.65 mm), paramere longer (ratios PL:HW > 0.20, PL:AEL > 1.15, PL:SH > 0.90) (Figs 1B, 7A–R) S. oahuensis sp. n.
Darker species (generally dark brown to black), fore wing vein Rs typically shorter relative to wing length (ratio WL:Rs 1.76–2.03) resulting in longer distance between terminations of Rs and M1+2, antennae shorter (AL ≤ 0.65 mm), paramere shorter (ratios PL:HW < 0.20, PL:AEL < 1.15, PL:SH < 0.90) (Figs 1C, 4A–T) S. atra sp. n.