Table 2. Summary of model parameters.

Parameter/variable	Description
nxy, rxy	number and growth rate of xy-type cells
β0	benefit from niche construction for cheaters and 2° producers
β1	benefit from niche construction for local and global producers
k	intrinsic carrying capacity
α	intravasation rate
θ, ϕ, ω, ψ, μ, ν	interclonal competition terms (see Fig 2)
g	resource production rate
l	independent resource depletion rate

A summary of the model parameters, some of which are estimated as described in S3 Appendix.