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Ethogram of behavior for the scan sampling methods
| Behavior categories | Description |
|---|---|
| Rest | Not moving any part of the body |
| Crawl | Alternate placement of feet or/and thumb claws to shift position on the rock face |
| Groom | Scratching or licking the body or/and wing phalanges |
| Fight | Biting or hooking (rapid swiping with thumb claw) the opponent |
| Consume ectoparasites | Taking ectoparasites into the mouth, masticating, and no evidence of rejecting them |
