Table 4.

Nucleotide substitution rate and selection pressures for the HAdV-55 major capsid protein genes.

Gene	Length(bp)	No. of sequences	Mean Non-synonymous Substations/site	Mean Synonymous Substitutions/site	dN/dS
Hexon	2839	23	0.00004	0.00064	0.06254
Penton base	1674	17	0.00048	0.00059	0.81874
Fiber	978	43	0.00037	0.00039	0.96781

The mean numbers of non-synonymous (dN) and synonymous(dS) substitutions per site from between sequences were noted and dN/dS were calculated. This selection pressure analysis was conducted using the maximum-composite-likelihood model, and included nucleotide sequences from 23 hexon, 43 fiber, and 17 penton base genes of HAdV-55 available from GenBank. All positions containing gaps were eliminated automatically. Evolutionary analyses were performed with MEGA 5.05 version.