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Evolution of deep-grazing herbivore groups since the Cambrian. The capacity to excavate calcareous coralline algae is taken as an indication of improved feeding capability. Left panel after Steneck 1983, Right photos include parrotfish bite marks, parrotfish, limpet bites (scale 0.5 mm) and sea urchin bites into calcareous algae (all photos by R. Steneck).
