Mean percentage of each strategy in the population varied based on level of skew in contact pattern (no skew, low skew, high skew), forgetting rate (λ specifies starting point, ψ specifies decay rate) and type of memory error (guessing or skipping). ALLD represents all defect strategy. TFT family sums the percentages from strategies tit-for-tat, tit-for-two-tats, generous tit-for-tat and contrite tit-for-tat. GRIM represents grim trigger strategy. Standard error of the mean error bars are not shown because they are smaller than the data point symbols. (Online version in colour.)