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. 2018 Jun 21;33(2):175–184. doi: 10.3803/EnM.2018.33.2.175

Table 2. Susceptibility Loci for Levels of Thyroid Stimulating Hormone or Free Thyroxine Detected by Genome-Wide Association Studies.

Phenotypes Locus Gene Protein function Population Reference
TSH 5q13.3 PDE8B Role in hydrolysis of the second messenger cAMP European, USA, Germany, UK [38,40,48,49]
6q27 PDE10A Role in regulation of the intracellular concentration of cyclic nucleotides European, UK [38,40]
1p36.13 CAPZB Regulating actin filament dynamics European, Chinese Han, Germany, UK [38,39,40,49]
16q23.2 MAF Role in increased T-cell susceptibility to apoptosis European, UK, Germany [38,40,49]
6p21.1 VEGFA Proliferation and migration of vascular endothelial cells European, UK [38,40]
4q31.23 NR3C2 Role in aldosterone actions European, Germany, UK [38,40,49]
2q35 IGFBP5 Encoding insulin like growth factor binding protein 5 European [38]
17q24.3 SOX9 Role in chondrocyte differentiation European [38]
1p31.3 NFIA Encoding nuclear factor IA European [38]
15q21.2 FGF7 Mitogenic and cell survival activities European [38]
11p11.2 PRDM11 Role in transcription regulation European [38]
15q26.1 MIR1179 MicroRNA 1179 European [38]
19p13.2 INSR Encoding insulin receptor European [38]
9q34.2 ABO Determination of ABO blood group European, UK [38,40]
14q32.12 ITPK1 Regulation of the synthesis of inositol tetraphosphate European [38]
8p12 NRG1 Role in the growth and development of multiple organ systems European [38]
14q13.3 MBIP-NKX2-1 Encoding TTF-1, binding to TG promoter European, UK [38,40]
6q24.3 SASH1 Role in the TLR4 signaling pathway European [38]
9p24.2 GLIS3 Role in transcription in thyroid gland European [38]
8q12.1 XKR4 Role in apoptosis Chinese Han [39]
9q22.33 FOXE1 Encoding TTF-2, role in thyroid morphogenesis Chinese Han, USA, UK [39,40,48]
2q35 IGFBP2 Encoding insulin like growth factor binding protein 2 UK [40]
3p25.2 SYN2 Binding to small synaptic vesicles UK [40]
Free T4 1p32.3 DIO1 Encoding iodothyronine deiodinase 1 European, UK [38,40]
9q34.3 LHX3 Role in pituitary development European, UK [38,40]
9q22.33 FOXE1 Encoding TTF-2, role in thyroid morphogenesis European [38]
4q33 AADAT Role in L-lysine catabolism European, UK [38,40]
16q12.2 LPCAT2-CAPNS2 Role in membrane biogenesis European [38]
18q22.3 NETO1-FBXO15 Role in spatial learning and memory in the hippocampus European [38]
18q12.1 B4GALT6 Role in biosynthesis of glycosphingolipids UK [40]

TSH, thyroid stimulating hormone; PDE8B, phosphodiesterase 8B; cAMP, cyclic adenosine monophosphate; PDE10A, phosphodiesterase 10A; CAPZB, capping actin protein of muscle Z-line subunit beta; VEGFA, vascular endothelial growth factor A; NR3C2, nuclear receptor subfamily 3 group C member 2; IGFBP5, insulin like growth factor binding protein 5; SOX9, SRY-box 9; NFIA, nuclear factor I A; FGF7, fibroblast growth factor 7; PRDM11, PR/SET domain 11; MIR1179, microRNA 1179; INSR, insulin receptor; ITPK1, inositol-tetrakisphosphate 1-kinase; NRG1, neuregulin 1; MBIP, MAP3K12 binding inhibitory protein 1; NKX2-1, NK2 homeobox 1; TTF, thyroid transcription factor; TG, thyroglobulin; SASH1, SAM and SH3 domain containing 1; TLR4, Toll-like receptor 4; GLIS3, GLIS family zinc finger 3; XKR4, XK related 4; FOXE1, forkhead box E1; IGFBP2, insulin like growth factor binding protein 2; SYN2, synapsin II; T4, thyroxine; DIO1, iodothyronine deiodinase 1; LHX3, LIM homeobox 3; AADAT, aminoadipate aminotransferase; LPCAT2, lysophosphatidylcholine acyltransferase 2; CAPNS2, calpain small subunit 2; NETO1, neuropilin and tolloid like 1; FBXO15, F-box protein 15; B4GALT6, beta-1,4-galactosyltransferase 6.