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. 2018 Jul 2;8:9961. doi: 10.1038/s41598-018-28177-4

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Mutability landscape of ADS for expression level using quantitative densitometric assay. For the mutability landscape, the vertical axis portrays the 20 possible amino acid residues. The wild-type amino acid residue at each position is indicated by bold squares, white squares represent variants that are not present in the library and grey squares represent variants that are not expressed. The color represents the concentration of expressed protein where blue range squares indicate decrease in expression while red range squares indicate increase in expression compared to the wild type.