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. 2018 Jul 9;8:10345. doi: 10.1038/s41598-018-28504-9

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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The study area (Iran) and patterns of endemism. (a) About half of the study area belongs to the Irano-Anatolian biodiversity hotspot (whitish area). Biogeographically, the study area belongs mainly to the Irano-Turanian region (dashed area), bounded by the Euro-Siberian and the Saharo-Sindian regions in the north and south, respectively. Major mountain ranges are indicated. (b) Endemic richness of Asteraceae per grid cell, only taking species within the study area into account, and identified AEs (outlined in colour).