Contribution of the storage effect () and relative nonlinearity () to the growth rate () of each species as an invader () in pairwise competition, where = average fitness differences + + . (A) Pairs where coexistence is predicted because for both species. In the case of M. koreensis (Mk) and M. reukaufii (Mr), would be negative for M. koreensis without the contribution of , and negative for M. reukaufii without the contribution of . In the case of S. bombicola (Sb) and M. gruessii (Mg), M. gruessii would still have without either mechanism, while persistence of S. bombicola requires one or both mechanisms. (B) Pairs where coexistence is not predicted because one species has . In all pairs, is weak to nonexistent, while is strongly equalizing (i.e., differences in would be much larger without the contribution of ).