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Conserved cysteines in L2 are required for WW1 binding. RG1 and WW1 binding with either wild-type (WT) HPV16 PsV or HPV16 point mutant PsV at the HPV16L218–32 site was assessed by Western blotting. The WW1 epitope on HPV16 and the corresponding L2 point mutant regions on other mutants were aligned using UCSF chimera software. “+” indicates strong binding, “+/−” indicates weak binding, and “−” indicates no binding.
