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. 2018 Jun 29;115(29):E6789–E6798. doi: 10.1073/pnas.1714044115

Table 3.

Variable definitions

Symbol Unit Description
Traits (constant through ontogeny)
ω kg Seed mass (SM)
Hmat m Height at maturation (Hmat)
ν kgm−2 Leaf nitrogen content per unit leaf area (N)
ϕ kgm−2 Leaf mass per unit leaf area (LMA)
ρ kgm−3 Wood density (WD)
State or rate variables (may vary through ontogeny)
H m Height of a plant
D m Stem diameter
Mi kg Mass of tissue type i retained on plant
Ai m2 Surface or cross-section are of tissue type i
dBdt kgy1 Biomass growth rate
p,p¯ moly1m2 Photosynthetic rate per unit area
ri moly1kg1 Respiration rate per unit mass of tissue type i
ki y1 Turnover rate for tissue type i
Environmental variables (fixed)
E Canopy openness
Other parameters (constant throughout)
αy Yield, fraction of carbon fixed into biomass
αbio kgmol−1 Biomass per mol carbon
ηc Crown-shape parameter
θ Sapwood area per unit leaf area
αl1 m Height of plant with leaf area of 1m2
αr1 kgm−2 Root mass per unit leaf area
αb1 Ratio of bark area to sapwood area

For Mi , ri , ki and Ai , subscripts refer to: a, alive tissue; b, bark; h, heartwood; l, leaves; r, roots; s, sapwood; st, total stem.