Maximum clade credibility tree inferred in BEAST2. A strict clock and coalescent Bayesian skyline population prior were used. The substitution rate is inferred as 1.22 × 10−5 s/s/y (95% HPD interval, 1.04 × 10−5–1.40 × 10−5 s/s/y), and the root age as 10.1 kya (95% HPD interval, 9.0–11.3 kya). The x axis denotes time into the past, in years. Taxon names: genotype/historical period, accession number/sample identifier, sample age, country abbreviation of sequence origin and region of sequence origin, as determined by the Standard country or area codes for statistical use (42). Horizontal gray bars indicate 95% HPD intervals of node ages.