Surface area of folded
regions of ovalbumin (Ova) estimated by
Congo red (CR) binding assay. (a) Visible absorbance of CR over a
range of CR:ovalbumin concentration from 0.8 to 27, neutral pH. CR
bound to ovalbumin secondary structure absorbs at a longer wavelength
than CR free in solution, so as ovalbumin is added to the solution
relative to CR, thereby increasing opportunities for CR to bind ovalbumin,
the overall absorption of the system shifts to the right. (b) Same
as part a, but in 0.5 M NaOH with the CR:Ova concentration from 0.8
to 14. A similar peak shift, of a smaller overall magnitude, was observed
at high pH. (c) CR absorbance peak as a function of CR:Ova concentration
ratio. Blue symbols label CR:Ova at neutral pH, and red symbols label
CR:Ova in high pH consistent with pidan formation. (d) Calculated
number of CR bound to one ovalbumin monomer as a function of CR:Ova
concentration ratios for both neural pH (blue) and high pH (red).
Because CR only binds to secondary structure but not random coil,
this assay allows us to estimate the surface area of secondary structure
in the native protein compared to that of the conformation found in
pidan. The results suggest that the pidan conformation consists of
some secondary structure, and that the surface area of secondary structure
is roughly 60% that of the native conformation.