As the spindle forms during meiosis I and II, acentric fragments also experience a transverse force that eliminates them to the spindle periphery even as they are moving poleward. (A–D) Mid-prometaphase II cell in which a fragment, originally located near the spindle equator along the pole-to-pole axis (vertical line in B–D), moved laterally toward the spindle periphery as it migrated poleward. (E–H) Similar to A–D, except this fragment (F–H, arrow) was created near the spindle pole during prometaphase I. The fragment was slowly translated laterally toward the spindle periphery, which is outlined by a sheath of mitochondria. Bar (in H), 5 μm.