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Reactions of 4-hydroxyphenylpyruvate oxygenases. (A) In vancomycin biosynthesis, HmaS-mediated hydroxylation affords L-4-hydroxymandelate 150. (B) HPPD hydroxylates aromatic carbon to yield homogentisate. (C) In the reaction catalyzed by the HPPD F337I variant, both 150 and 154 are produced.236
