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The relationship between model quality and fit to data breaks down in deposited 3.5–4 Å resolution crystal (a) and cryo-EM (b) models. The crystal structure cohort was limited to models deposited from 2007 to the present, and R free values for data at ≤3.75 Å and >3.75 Å resolution are offset by ±0.0015 for clarity. All EM models in the resolution range with masked CC ≥ 0.5 were included in the analysis.
