. Author manuscript; available in PMC: 2019 Jan 9.
Published in final edited form as: Nat Genet. 2018 Jul 9;50(8):1140–1150. doi: 10.1038/s41588-018-0156-2
Users may view, print, copy, and download text and data-mine the content in such documents, for the purposes of academic research, subject always to the full Conditions of use: http://www.nature.com/authors/editorial_policies/license.html#terms