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. 2018 Aug 17;8:12347. doi: 10.1038/s41598-018-30646-9

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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The state model of chromatin in 1q21.3 (A) and 20q13.12 (B). The chromatin was segmented into 25 states (i.e., promoter, enhancer) with the ChromHMM and ChromImpute algorithms^45,46 by using data from the NIH Roadmap Epigenomics Consortium (http://www.roadmapepigenomics.org) and was visualized with the WashU Epigenome Browser (http://epigenomegateway.wustl.edu/browser/). Here, 25 states are summarized to 9 as shown in the color legend. The gray shaded stripe represents an active/weak enhancer (A) or bivalent promoter regions (B).