Net diversification (birth − death, r = b − d, events per million years) estimated from known standing diversity and crown or stem ages for all diatoms (ON + AN + AM), the bilaterally symmetrical clade (pennate diatoms, AN + AM) and the actively motile clade (raphid diatoms, AM). Shown are the mean values (circles) and minimum–maximum range (error bars) across bootstrap phylogenies. All calculations used relative extinction ε = 0.751 estimated from Cenozoic fossil data (Table 1). Numbers next to the points indicate the total number of species in the respective databases. Numbers below the clade names are the estimates of crown and stem ages in million years ago (Ma, Fig. 1). ON, oogamous nonmotile; AN, anisogamous nonmotile; AM, anisogamous motile.